RT Journal Article T1 Human calcanei from the Middle Pleistocene site of Sima de los Huesos (Sierra de Atapuerca, Burgos, Spain) A1 Pablos, Adrián A1 Martínez Mendizábal, Ignacio A1 Lorenzo, Carlos A1 Sala Burgos, Nohemi A1 Gracia Téllez, Ana A1 Arsuaga Ferreras, Juan Luis AB The existence of calcanei in the fossil record prior to modern humans and Neandertals is very scarce. This skeletal element is fundamental to understanding the evolution of the morphology of the foot in human evolution. Here we present and metrically and comparatively describe 29 calcaneus remains from the Middle Pleistocene site of Sima de los Huesos (SH) (Sierra de Atapuerca, Burgos, Spain). These calcanei belong to 15 individuals (nine adults, two adolescents and four immature individuals). The metric and morphological differences in the calcanei among Middle and Late Pleistocene hominins tend to be subtle. However, the calcanei from SH are broad and robust with large articular surfaces and most significantly, exhibit a very projected sustentaculum tali. A biomechanical and phylogenetic interpretation is proffered to explain the observed morphology of these calcanei. It has been possible to propose tentative sex assignments for the SH calcanei based on size, using methods similar to those used to establish sex from the talus bones from SH. The estimation of stature based on the calcaneus provides a mean of 175.3 cm for males and 160.6 for females, which is similar to that obtained using other skeletal parts from the site. In sum, the SH calcanei are robust with a proportionally long tubercle and a projected sustentaculum tali, which are traits shared by Neandertals. PB Elsevier SN 0047-2484 YR 2014 FD 2014-11 LK https://hdl.handle.net/20.500.14352/34308 UL https://hdl.handle.net/20.500.14352/34308 LA eng NO Unión Europea. FP7 NO Ministerio de Ciencia e Innovación (MICINN) NO Ministerio de Economía y Competitividad (MINECO) NO Generalitat de Catalunya NO Fundación Atapuerca NO Junta de Castilla y León DS Docta Complutense RD 8 abr 2025