RT Journal Article T1 The carnivore remains from the Sima de los Huesos Middle Pleistocene site (Sierra de Atapuerca, Spain) A1 García García, Nuria A1 Arsuaga Ferreras, Juan Luis A1 Torres, T. AB Remains of carnivores from the Sima de los Huesos site representing at least 158 adult individuals of a primitive (i.e., not very speleoid) form of Ursus deningeri Von Reichenau 1906, have been recovered through the 1995 field season. These new finds extend our knowledge of this group in the Sierra de Atapuerca Middle Pleistocene. Material previously classified as Cuoninae indet. is now assigned to Canis lupus and a third metatarsal assigned in 1987 to Panthera cf. gombasz oegensis, is in our opinion only attributable to Panthera sp. The family Mustelidae is added to the faunal list and includes Martes sp. and a smaller species. The presence of Panthera leo cf. fossilis, Lynx pardina spelaea and Felis silvestris, is confirmed. The presence of a not very speloid Ursus deningeri, together with the rest of the carnivore assemblage, points to a not very late Middle Pleistocene age, i.e., oxygen isotope stage 7 or older. Relative frequencies of skeletal elements for the bear and fox samples are without major biases. The age structure of the bear sample, based on dental wear stages, does not follow the typical hibernation mortality profile and resembles a catastrophic profile. The site was not a natal or refuge den. The hypothesis that the site was a natural trap is the most plausible. If the Sima de los Huesos functioned as a natural trap (without an egress out), the human accumulation cannot be attributed to carnivore activities and must be explained differently. PB Elsevier SN 0047-2484 YR 1997 FD 1997-08 LK https://hdl.handle.net/20.500.14352/122581 UL https://hdl.handle.net/20.500.14352/122581 LA eng NO Garcı́a, N., Arsuaga, J. L., & Torres, T. (1997). The carnivore remains from the Sima de los Huesos Middle Pleistocene site (Sierra de Atapuerca, Spain). Journal of Human Evolution, 33(2-3), 155-174. https://doi.org/10.1006/jhev.1997.0154 NO Dirección General de Investigación Científica y Técnica (España) NO Junta de Castilla y León DS Docta Complutense RD 18 jul 2025