New dinosaur sites correlated with Upper Maastrichtian pelagic deposits in the Spanish Pyrenees: implications for the dinosaur extinction pattern in Europe

Abstract

Six new dinosaurs sites have been found close to the Cretaceous/Tertiary boundary in Arén (south-central Pyrenees, Huesca, Spain) in coastal and non-marine deposits of the Arén and Tremp Formations. The sites contain articulated remains (skull elements, vertebrae, hind-limb bones) and isolated teeth and bones of hadrosaurids, three types of theropod teeth, one sauropod, at least seven types of eggshells (six ornithoid types and one Megaloolithidae probably from a sauropod), remains of other vertebrates, and four charophyte species. The fossil-bearing rocks have been correlated with marine sediments containing planktonic foraminifera from the uppermost Maastrichtian Abathomphalus mayaroensis Biozone. These rich and diversified dinosaur assemblages enable more accurate dating of the faunal changes that took place during the Maastrichtian in Europe and support the hypothesis of a sudden dinosaur extinction at the Cretaceous/Tertiary boundary.