On the Occurrence of the Gar Obaichthys africanus Grande in the Cretaceous of Portugal: Palaeoecological and Palaeobiogeographical Implications

Abstract

Several disarticulated fish scales and a vertebral centrum were collected from the uppermost middle to lowermost upper Cenomanian beds of the vertebrate fossil site of Casais dos Carecos (Baixo Mondego region, Coimbra, West Portugal), corresponding to shallow carbonate and mixed facies (“Unit B”) of the Tentúgal Formation. The taphonomic preservation of these remains is adequate for their detailed taxonomical study. The scales are diagnosable for the species Obaichthys africanus Grande, as each one has a strong spine protruding from the middle of the posterior border in a rostrocaudal direction, a dorsoventral peg and socket, and a small anterodorsal process for articulation of adjacent scales. The sculpture with ganoid ridges and grooves distinguishes the studied remains from those of O. decoratus Wenz and Brito, which display rounded tubercles. The isolated Lepisosteoidea centrum is tentatively attributed to O. africanus as well, as it was found together with the scales, and no other compatible remains were found in the site. The fish material reported here constitutes the first record of O. africanus in the Baixo Mondego region and northern ranges of the Albian–Turonian West Portuguese Carbonate Platform, as well as in the uppermost middle to basal upper Cenomanian of Portugal.