Reworked marine pythonomorph (Reptilia, Squamata) remains in Late Pleistocene cave deposits in central Spain

Abstract

Three sites at the Calvero de la Higuera complex (Camino Cave, Navalmaíllo Rock Shelter, and Buena Pinta Cave), near the village of Pinilla del Valle (Madrid, Spain), are known for their record of Neanderthals and other Late Pleistocene mammals. Occasionally, they also yield much more ancient reworked remains, which come from the Upper Cretaceous dolomites and carbonatic sandstones in which these caves and shelters are developed. These are mostly teeth of sharks and bays and vertebrae and teeth of bony fishes, but several reptile vertebrae and teeth have also been found. These reptile remains, which we describe here, likely belong to the pythonomorph incertae sedis Carentonosaurus cf. mineaui. This taxon is known from several outcrops in the southwest of France but is rare on the Iberian Peninsula; indeed, only a few remains possibly related to the genus have been found (in the Caba~na Formation, Asturias, Spain). The pythonomorph remains discussed here are the first fossils of marine reptiles from the Madrid region. Should the assignment to Carentonosaurus be confirmed, the teeth would provide novel data on the characteristics of this rather poorly known taxon, and might help clarify its phylogenetic relationship within Pythonomorpha.